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Gijs Kronenberg

Gijs C. Kronenberg (*1957) started looking at strombs about 30 years ago. This lead to the publication together with the late Jan Berkhout of a popular review in Dutch of Strombidae in the loose-leaf edition of Vita Marina (Kronenberg & Berkhout, 1984). After a few more papers on Stromboidea, focus changed to other families of gastropods, including Xenophoridae (which may turn out to be Stromboideans, as suggested by L.R.L. Simone, 2005 and S. Kiel & M. del Carmen Perrilliat, 2001) Personidae and Columbellidae, in the supposition that taxonomy of Stromboidea was clearified.

However, after restudying specimens it turned out that this is not the case. This has resulted in an ongoing study of especially Strombidae and Rostellariidae (the latter recognised as a family closely related, but distinguished from Strombidae). Studies are undertaken partially alone, partially in cooperation with others, both malacologists and palaeontologists.

Main focuses of interest are taxonomy and phylogeny of Strombidae and Rostellariidae.

He has served in the editorial board of Vita Marina from Vol. 41 (1991) onwards till it stopped in 2001. He then joined the editorial staff of it's "successor" Vita Malacologica from No1. onwards, and starting with Vol. 73 he has joined the editorial staff of Basteria, the scientific journal of the Netherlands Malacological Society.

List of Publications on Stromboidea, including Xenophoridae, by Gijs C. Kronenberg

  • 1984: [dated 1981] Kronenberg, G.C. & Berkhout, J.: Strombidae. – Vita Marina 31(1-6): buikpotigen 263-362, pls. 1-9.
  • 1986a: Kronenberg, G.C. & Berkhout, J.: Strombidae II: addenda & corrigenda. – Vita Marina 36(1-3): buikpotigen 363-368.
  • 1986b: Kronenberg, G.C.: An extension of the range of Strombus (Dolomena) plicatus plicatus (Röding, 1789) (Gastropoda: Strombidae). – Basteria 50(4-6):151-154.
  • 1988a: Kronenberg, G.C.: Strombus (Canarium) cf. ochroglottis Abbott, 1960 van Sri Lanka. – C.B. Ned. Malac. Ver. 240: 388-389.
  • 1988b: Kronenberg, G.C. & Goud, J.: Xenophoridae, een overzicht van alle recent voorkomende soorten (I). – Vita Marina 38(4-6): buikpotigen 457-480
  • 1989: Kronenberg, G.C. & Goud, J.: Xenophoridae, een overzicht van alle recent voorkomende soorten (II). – Vita Marina 39(1-2): buikpotigen 481-494.
  • 1991: Kronenberg, G.C.: Strombus (Canarium) ochroglottis betuleti, a new subspecies from Sri Lanka with a short note on the distribution of S. (C.) mutabilis Swainson, 1821. – Gloria Maris 30(4): 53-58.
  • 1992: Kronenberg, G.C.: The Recent species of the family Aporrhaidae. – Vita Marina 41(2): 73-84.
  • 1993: Kronenberg, G.C.: On the identity of Lambis wheelwrighti Greene, 1978 and L. arachnoides Shikama, 1971. – Vita Marina 42(2): 41-56.
  • 1997: Kronenberg, G.C.& Kamphuis, A.: Does Strombus peruvianus Swainson, 1823, live off the coast of Baja California Sur? – The Festivus 29(2): 14.
  • 1998: Kronenberg, G.C.: Revision of Euprotomus Gill, 1870. 1. The systematic position of Strombus listeri Gray, 1852 (Gastropoda Prosobranchia: Strombidae). – Vita Marina 45(3-4): 1-6.
  • 1999a: Kronenberg, G.C.: Revision of Euprotomus Gill, 1870. 2. On the identity of Strombus hirasei Kuroda, 1942 (Gastropoda Prosobranchia: Strombidae). – The Festivus 31(6): 63-67.
  • 1999b: Kronenberg, G.C.: Enkele opmerkingen met betrekking tot de Strombus-soort uit de Middellandse Zee. – Correspondentieblad van de Nederlandse Malacologische Vereniging 309: 91-93.
  • 1999c: Kreipl, K., Alf , A. & Kronenberg G.C. 1999. A new subgenus of the family Xenophoridae (Mollusca, Gastropoda). – Spixiana 22(2): 179-180.
  • 2000a: Kronenberg, G.C.: & Dekker, H.: A nomenclatural note on Strombus wilsoni Abbott, 1967, and Strombus wilsonorum Petuch, 1994, with the introduction of Strombus praeraninus nom. nov. (Gastropoda, Strombidae). – Basteria 64 (1-3): 5-6.
  • 2000b: Kronenberg, G.C.: Strombus gigas bewerkt tot steekwapen: bestaat dat echt? – Correspondentieblad van de Nederlandse Malacologische Vereniging 317: 145-147.
  • 2002a: Kronenberg, G.C.: Addendum to a revision of Euprotomus Gill, 1870. 2a. On the identity of Strombus hirasei Kuroda, 1942 (Gastropoda Prosobranchia: Strombidae). – The Festivus 34(9): 110.
  • 2002b: Kronenberg, G.C. & Burger, A.W.: On the subdivision of recent Tibia-like gastropods (Gastropoda: Stromboidea) with the recognition of the family Rostellariidae, Gabb, 1868 and a note on the spelling of a species. – Vita Malacologica 1: 39-48.
  • 2002c: Kronenberg, G.C. & Vermeij, G.J.: Terestrombus and Tridentarius, new genera of Indo-Pacific Strombidae (Gastropoda), with comments on included taxa and on shell characters in Strombidae. – Vita Malacologica 1: 49-54
  • 2002d: Kronenberg, G.C.: Revision of Euprotomus Gill, 1870. 3. Description of Euprotomus aurora spec. nov. from the Indian Ocean (Gastropoda, Strombidae). – Vita Malacologica 1: 55-60.
  • 2004: Willan, R.C. & Kronenberg, G.C.: Rectification of nomenclature for two species in the family Strombidae (Gastropoda). – Basteria 67(4-6): 153-157.
  • 2005a: Kronenberg, G.C. & Lee, H.G.: Strombus granulatus Swainson, 1822 (Gastropoda: Strombidae), a very variable species, including a note on homonymy with Strombus granulatus Röding, 1798 (Gastropoda: Cerithiidae). – The Festivus 37(3): 31-35.
  • 2005b: Kronenberg, G.C.: Nogmaals Strombus gigas bewerkt tot steekwapen. – Spirula 343: 57.
  • 2005c: : Kronenberg, G.C. & Dharma, B.: New distributional records for four species of Stromboidea (Mollusca: Gastropoda) from Australasia. – The Beagle, Records of the Museums and Art Galleries of the Northern Territory 21: 47-51.
  • 2006: Burger, A.W. & Kronenberg, G.C.: The occurrence of Ectinochilus Cossmann, 1889 (Gastropoda; Strombidae) in New Zealand, with the description of two new Eocene species. – Molluscan Research 26(2): 77-83.
  • 2007: Kronenberg, G.C. & Lee, H.G.: Genera of American strombid gastropods (Gastropoda: Strombidae) and remarks on their phylogeny. – The Veliger 49(4): 256-264.
  • 2008a: Kronenberg, G.C.: Born’s strombs (Mollusca: Gastropoda), with some notes on Strombus succinctus Linnaeus, 1767. – Annalen des Naturhistorischen Museums in Wien, Serie B, 109: 51 – 66.
  • 2008b: : Landau, B., Kronenberg, G.C. & Herbert, G.: A large new species of Lobatus (Gastropoda: Strombidae) from the Neogene of the Dominican Republic, with notes on the genus. – The Veliger 50(1): 31-38.
  • 2008c: Kronenberg, G.C.: Family Rostellariidae. [in]. Poppe, G. [ed.]. Philippine marine mollusks. I, Gastropoda 1: 512, pl. 201. Conchbooks, Hackenheim, Germany.
  • 2008d: Kronenberg, G.C.:Family Strombidae. [in]. Poppe, G. [ed.]. Philippine marine mollusks. I, Gastropoda 1: 514-570, pls 202-230. Conchbooks, Hackenheim, Germany.
  • 2008e: Kronenberg, G.C.:Family Seraphsidae. [in]. Poppe, G. [ed.]. Philippine marine mollusks. I, Gastropoda 1: 572, pl. 231. Conchbooks, Hackenheim, Germany.
  • 2008f: Harzhauser, M. & Kronenberg, G.C.: A Note on Strombus coronatus Defrance, 1827 and Strombus coronatus Röding, 1798 (Mollusca: Gastropoda). – The Veliger 50(2): 120-128.
  • 2008g: Kronenberg, G.C.: An intergeneric hybrid (Gastropoda: Caenogastropoda: Strombidae) with remarks on the subdivision of Indo-Pacific Tricornis. – Basteria 72 (4-6): 331-343.
  • 2009a: Kronenberg, G.C., Liverani, V. & Dekker, H.: On the identity of Strombus coniformis Sowerby II, 1842 (Gastropoda, Strombidae) with additional notes on its distribution. – Journal of Conchology 39(6): 659-668.
  • 2009b. Kronenberg, G.C.: A note on Bandel's 2007 review of the classification and phylogeny of the Strombidae (Caenogastropoda). – Basteria 73(1-3): 65-67.
  • 2010: Landau B.M., Kronenberg, G.C., & da Silva, C.M. A new species of Lobatus (Mollusca: Caenogastropoda: Strombidae) from the Neogene of the Dominican Republic, with notes on further species from the Dominican assemblages. – Basteria 74(4-6): 95-109.
  • 2011a: Landau B.M., Kronenberg, G.C., Herbert, G.H., & da Silva, C.M.: The genus Strombus (Mollusca: Caenogastropoda: Strombidae) in the Neogene of the Bocas del Toro area, Panama, with the description of three new species. – Journal of Palaeontology 85(2): 337-352.
  • 2011b: Kronenberg, G.C.: Lobatus gigas (Linnaeus, 1758) (Gastropoda, Strombidae) in Shark Bay? – Spirula 379: 37.
  • 2011c: Kronenberg, G.C.:Family Strombidae Rafinesque, 1815 – review and comments. [In] Severns, M. [ed.] Shells of the Hawaiian Islands - the Sea Shells 128-131, pls 48-49. Conchbooks, Hackenheim, Germany 1-562.
  • 2012a: Kronenberg, G.C.: Gibberulus gibbosus (Röding, 1798) (Caenogastropoda: Strombidae) from the Islas Galápagos? – the Festivus 44( 4): 46-47.
  • 2012b: Kronenberg, G.C. & Harzhauser, M.: Europrotomus (Mollusca: Caenogastropoda: Strombidae): a new Middle Miocene European strombid genus (Revision of Euprotomus Gill, 1870. Part 4). – Paläontologische Zeitschrift 86(2): 147-159. DOI 10.1007/s12542-011-0121-1.
  • 2012c: Kronenberg, G.C.: Identifications for Stromboidea (Caenogastropoda) in the facsimile reprint of the Favanne de Montcervelle 1780 edition of “La Conchyliologie, ou Histoire naturelle …” by Dézallier d’Argenville. – Conchylia 42(1-4): 3-12.

papers accepted:

  • Harzhauser, M. & Kronenberg, G.C.: The Neogene Persististrombus-lineage (Gastropoda: Strombidae) in the Paratethys Sea. – Acta Palaeontologica Polonica

Strombid taxa described:

  • Strombus (Canarium) ochroglottis betuleti (1991) named after Jan Berkout, now considered *Canarium betuleti
  • Mirabilistrombus (1998)
  • Strombus praeraninus (2000) as a new name for Strombus wilsonorum Petuch, 1994 non Abbott, 1967 together with Henk Dekker.
  • Terestrombus (2002) with Geerat Vermeij
  • Tridentarius (2002) with Geerat Vermeij
  • Euprotomus aurora (2002)
  • Ectinochilus trisfragmentum (2006) with Adri Burger
  • Ectinochilus antipodarum (2006) with Adri Burger
  • Persististrombus (2007) with Harry G. Lee
  • Lobatus vokesae Landau, Kronenberg & Herbert, 2008
  • Lobatus pascaleae Landau, Kronenberg & da Silva, 2010
  • Strombus acanthospira Landau, Kronenberg, Herbert & Silva, 2011
  • Strombus elegantissimus Landau, Kronenberg, Herbert & Silva, 2011
  • Strombus vermeiji Landau, Kronenberg, Herbert & Silva, 2011
  • Europrotomus Kronenberg & Harzhauser, 2012

Examples from Gijs Kronenbergs collection:

Dolomena kleckhamae (Cernohorsky, 1971); East of Hannover Island, Papua New Guinea; dredged alive at 110m on rubble; Coll. G.C. Kronenberg No. 6302

Dolomena kleckhamae (Cernohorsky, 1971); East of Hannover Island, Papua New Guinea; dredged alive at 110m on rubble; Coll. G.C. Kronenberg No. 6302

Dolomena plicata; Ras Zafarana, Gulf of Suez, Egypt; 9/1992; Coll. Gijs Kronenberg No. 6323

Dolomena plicata; Sharm el Naga, Egypt, Red Sea; dead on beach, 70,7 mm; Coll. Kronenberg No. 5496

Strombus (Dolomena) plicatus plicatus; Ras Matarma, Gulg of Suez, Sinai, Egypt; dead on beach; 1969; Coll. Gijs Kronenberg No. 713

Dolomena plicata; Red Sea; Coll. Kronenberg No. 5497

Strombus blanci Paratype; Niau Atoll, Tuomoto Archipelago, French Polynesia; ~120.000 YBP, Pleistocene; Coll. Gijs Kronenberg No. 6387

Strombus blanci; Niau Atoll, Tuomoto Archipelago, French Polynesia; ~120.000 YBP, Pleistocene; Coll. Gijs Kronenberg No. 6369

Conomurex fasciatus; Ras Gemsa, west coast Gulf of Suez, 50 km north of Hurghada, 27° 39´ N, 33° 36´ E, Egypt; muddy flat merging into a sandy flat, both with stones and old coral blocks; 1988; Coll. Gijs Kronenberg No. 1480

Strombus (Conomurex) fasciatus; Port Sudan, Egypt; Coll. Gijs Kronenberg No. 1262

Tricornis taurus; Hospital Point near Tumon Bay, Guam; among silty sandy grass at 45-55 ft.; 1995; Coll. Gijs Kronenberg No. 6101

Strombus (Tricornis) taurus; Rongelap Atoll, Marshall Islands; inside lagoon on sand at 5-8 ft.; 3/1985; Coll. Gijs Kronenberg No. 5549

Sinustrombus latissimus x Lambis lambis hybrid; Vietnam, Binh Thuan province, approx. 250 km south of Nha Tranh; Summer 2007, by fishermen, at 15-20 m; Coll. Kronenberg No. 6331

Sinustrombus latissimus x Lambis lambis hybrid; Vietnam, off Nha Trang; 5/2008; 170 mm; Coll. Kronenberg No. 6331

Harpago chiragra x Lambis lambis; Vietnam, off Nha Trang; 2008; 230 mm; coll. Kronenberg No. 6366

Harpago chiragra; south of Sri Lanka; at 4-8 m on sand and rocks; 12/1979; Coll. Gijs Kronenberg No. 499

Lambis millepeda x truncata sebae; Cebu, Philippines; Coll. Kronenberg No. 5784

  • figured in Vita Marina 42(2):53, fig. 7, left

Lambis millepeda x truncata sebae; Sulu Island, Philippines; Coll. Kronenberg No. 5783

  • figured in Vita Marina 42(2):53, fig. 7, right

Lambis millepeda x truncata sebae; Malampaya, Palawan, Philippines; Coll. Kronenberg No. 2509

  • figured in Vita Marina 42(2):47, pl. 2, fig. 2a, 2b

Lambis arachnoides; Philippines; 180 mm; Coll. Kronenberg No. 439

  • figured in Vita Marina 42(2):49, pl. 3, fig. 2a, 2b

Lambis arachnoides; Bohol, Philippines; Coll. Kronenberg No. 440

Lambis millepeda x truncata; Cebu, Bantayan Island, Philippines; Co.. Gijs Kronenberg No. 6305

Lambis crocata x scorpius; Siquijar Island, Cebu, Philippines; Coll. Kronenberg No. 5117

Lambis crocata crocata; Nha Trang, Vietnam; 202 mm; 3/2007; Coll. Gijs Kronenberg, No. 6352A

  • Comment Gijs Kronenberg: "I have reached the preliminary conclusion that the specimen in question is not L. c. pilsbryi, but only a large L. c. crocata (Link, 1807)."

Lambis crocata pilsbryi; Marquesas Islands, Coll. Gijs Kronenberg No. 5767

Lambis crocata pilsbryi; Marquesas Islands, Coll. Gijs Kronenberg No. 875

Lambis violacea freak; Saint Brandon, Mauritius; Coll. Gijs Kronenberg No. 1835

Lobatus peruvianus; Pedro Gonzales Islands, Panama; Coll. Kronenberg, No. 7

Lobatus peruvianus; Panama; Coll. Kronenberg No. 702

Lobatus peruvianus; Baja California, Mexico; trawled at 10-15 m; Coll. Gijs Kronenberg No. 6341

Strombus (Tricornis) peruvianis; Medidora Island, Panama; dredged at 120 ft.; 1989; Coll. Gijs Kronenberg No. 1944

Lobatus haitensis; Gurabo Formation, Miocene; Dominican Republic; Coll. Gijs Kronenberg No. 6343

Lobatus williamsi; late Pliocene; Lee County, Florida, USA; Coll. Gijs Kronenberg No. 6335

Lobatus goliath; Fortaleza, Ceará, Brazil; 325 mm; Coll. Gijs Kronenberg No. 441

Lobatus goliath operculum; Fortaleza, Ceará, Brazil; 325 mm; Coll. Gijs Kronenberg No. 441

Lobatus goliath; Brazil; Coll. Gijs Kronenberg No. 637

Strombus gigas; Centent Key, Florida, USA; Coll. Gijs Kronenberg No. 328

Persististrombus latus; Senegal; Coll. Kronenberg No. 5404

Persististrombus latus; Senegal; Coll. Kronenberg No. 5405

Persististrombus granulatus; Isla del Cocos, Panama; by local diver; 36,6 mm; Coll. Gijs Kronenberg No. 3451

Strombus (Persististrombus) granulatus; Bahia de Chatham, Isla del Coco, Costa Rica; 3/1988; dredged at 80-100 m; leg. Hank Chaney et al., ex Coll. Harry G. Lee, Coll. Gijs Kronenberg No. 6299

  • Comment: This specimen is very similar to some forms of Persististrombus latus

Persististrombus granulatus; Isla Santa Fe, Islas Galapagos, Ecuador; taken in sand at 30 m, by local lobster diver, 2003; Coll. Gijs Kronenberg No. 6268

Persististrombus granulatus immature; Isla Santa Fe, Islas Galapagos, Ecuador; taken in sand at 30 m, by local lobster diver, 2003; Coll. Gijs Kronenberg No. 6268

Strombus bifrons; TU1215, Dominican Republic; Gurabo Formation; Coll. Gijs Kronenberg No. 6344

Mirabilistrombus listeri; Arafura Sea, between Aru and Australia; trawled by Indonesian fishermen at approx. 144 m; 2005; 116,8 mm; ex Coll. Bunjamin Dharma, Coll. Kronenberg No. 6289

Doxander japonicus; Fukuma, Fukuma Ken, Japan; Coll. Gijs Kronenberg No. 3072

Doxander japonicus; Mikawa, Aichi Pref., Japan; dredged at 5-10 fathoms; 1982; Coll. Gijs Kronenberg No. 5632

Doxander vittatus vittatus; Cooktown, Queensland, Australia; trawled offshore at 120 m; Coll. Gijs Kronenberg No. 110

Doxander vittatus vittatus; off Dingo Beach, Queensland, Australia; trawled; 1982; Coll. Gijs Kronenberg No. 528

Strombus (Canarium) maculatus depauperatus Dautzenberg & Bouge, 1933; Easter Islands, Vinapu; at 50 ft. on sand & Algae, by scuba diving; 5/1986; Coll. Kronenberg No. 5720

Strombus maculatus; Anakaena, Easter Islands; at 2 m. in sand around weeds; 10/1988; Coll. Gijs Kronenberg No. 2343

Strombus urceus orrae; Broome, Australia; Coll. Gijs Kronenberg No. 5676

Canarium urceus orrae; Monte Belo Islands, WA, Australia; 1982; Coll. Gijs Kronenberg No. 538

Strombus urceus orrae; Drimmie Head, NT, Australia; at low tide on sand flats; Coll. Gijs Kronenberg No. 5924

Canarium urceus incisus; Marau Sound, Guadalcanal, Solomon Islands; by diver at 30 m at night; Coll. Gijs Kronenberg No. 653

Canarium urceus incisus; Mele beach, Efate Island, Vanuatu; snorkeling at 3' to 5', dead; 8/1999; Coll. Gijs Kronenberg No. 598

Canarium urceus incisus; Langa Langa Lagoon, Malaita, Solomon Islands; Coll. Gijs Kronenberg No. 515

Strombus (Canarium) mutabilis ostergaardi Pilsbry, 1921; Kaneohe, Oahu, Hawaii, USA; Kaneohe sewer trench excavation, imbedded in hard pack matrix of sand / Limestone walls of trench blocks; 10/1994; Coll. Gijs Kronenberg No. 5130

Canarium klineorum; Trincomallee, Sri Lanka; 8/1983; Coll. Gijs Kronenberg No. 655

Tricornis tricornis; Red Sea, Egypt, West Coast Gulf of Suez, 10 km south of Safaga (26° 39´ 30´´ N, 33° 56´ 30´´ E); large muddy / sandy flat with coral blocks; 8/1988, Coll. Gijs Kronenberg No. 1457

Tricornis tricornis; Somalia; 1991; Coll. Gijs Kronenberg No. 2579

Tricornis tricornis immature; Somalia; 1991; Coll. Gijs Kronenberg No. 2579

Tricornis maximus (Martin, 1883), juvenile; Nyalindung, Western Java, Indonesia; Miocene; Coll. Gijs Kronenberg

Rostellariella lorenzi, paratype 5; Arafura Sea, north of Darwin, NT, Australia; trawled on mud and sand at 200-400 m by indonesian fishermen; 2004; exColl. Morrison, Coll. Kronenberg No. 6269

Rostellariella lorenzi; Arafura Sea, north of Darwin, NT, Australia; trawled on mud and sand at 200-400 m by indonesian fishermen; 2004; exColl. Morrison, Coll. Kronenberg No. 6294

Rostellariella barbieri Paratype 2; Oslob, south east Cebu, Philippines; with tangle nets at 200-300 m; 2007; Coll. Gijs Kronenberg No. 6324

Rostellariella delicatula;

  • top row:
    • left India off Madras, 1988, dredged, Coll. Kronenberg No. 1538;
    • middle India, off Madras, dredged, Coll. Kronenberg No. 0395;
    • right Andaman Sea, Coll. Kronenberg No. 5710
  • bottom row:
    • left Bangla Desh, Bay of Bengal, trawled, Coll. Kronenberg No. 5709;
    • Thailand, Coll. Kronenberg No. 0084;
    • Andaman Sea, Coll. Kronenberg No. 5711;
    • right Andaman Sea, Coll. Kronenberg No. 5711

Aporrhais uttingeriana; Pano Deftera, Cyprus; Pliocene; 1994; Coll. Gijs Kronenberg No. 6235

Aporrhais cf. triangulata; Herne Bay, Great Britain; Oldhaven Beds, Ypresian; Coll. Gijs Kronenberg No. 6236

Strombolaria crucis (Bayan, 1870); Albanello quarry, Chiampo, Vicenza, Italy; lower Lutetian, Eocene; Coll. Gijs Kronenberg No, 6242

Perissoptera schlotheimii; "Vijlen kalk", Maastrichtian; CBR quarry, Lixhe, Belgium; Coll. Kronenberg No. 6301

Perissoptera schlotheimii; "Vijlen kalk", Maastrichtian; CBR quarry, Lixhe, Belgium; Coll. Kronenberg

Aporrhais limburgensis; Enclosure 62A-13, Quarry Curfs, Geulhem, Netherlands; Maastrichtian, Maastricht-Formation, Meersen Member; Coll. Gijs Kronenberg No. 3258

Hippochrenes nuda (Binkhorst, 1861); Quarry ENCI, Meersen, Netherlands; ("Kalksteen van Meersen") Meersen Member, Maastricht-Formation, Maastrichtian; Coll. Gijs Kronenberg No. 6307

Calyptraphorus sp.; Mahajanpa(?), Madagascar; Maastrichtian(?), Cretaceous; Coll. Gijs Kronenberg No. 6232

from Gijs Kronenberg's photo collection:

Strombus hirasei Holotype; Photo Ron Voskuil, Copyright "The Festivus", Original published in Kronenberg, 1999

Varicospira sp; age unknown, probably Pliocene; Sibolga, Sumatera, Indonesia; 18,3 mm; Coll. ZMA, photo Ron Voskuil

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