/ Persististrombus Bernielandaui
- " Small-sized, thick-shelled, stout strombid with a row of knobby to spiny nodes on the teleoconch whorls and a row of 6-8 spines on the bodywhorl. Spiral threads cover the ramp of the angulated whorls; coarse spiral ribs appear on the bodywhorl which bears two irregular spirals of nodes or knobs. Thick, triangular pointed wing, demarcated from the bodywhorl by a deep concavity."
Original description of Strombus bernielandaui by Harzhauser, 2007:
- " Squat, robust, small-sized shell with high spire of 7 teleoconch whorls and broad conical bodywhorl; protoconch unknown. Teleoconch whorls weakly angulated in the lower third, resulting from a row of 7-9 knobs or nodes, which rarely develop spiny tips. Upper ramp covered by spiral threads, being best developed close to the irregular, wavy suture. On the bodywhorl, this delicate spiral sculpture is accompanied by much coarser, irregular spiral ribs. One of these ribs, in the middle of the bodywhorl, and a second one in the lower part, close to the stromboid notch, may develop knobs or spirally elongated nodes. The marked angulation of the bodywhorl coincides with a row of nodes or spines of variable number (6-8). Thus, the sculpture of the bodywhorl is very variable. The angulation persists into the wing, forming the tip of the pointed triangular wing. A deep concavity, below the last spine of the bodywhorl, separates the wing, which develops only the secondary spiral elements. The upper margin of the wing is straight; stromboid notch narrow and deep."
Locus typicus: Gebel Madrakah, Oman
Stratum typicum: Acropora bearing bioclastic limestones of the Warak Formation, Chattian, Oligocene
Holotype: NHMW 2006z0231/0035; width 39 mm, height 52.5 mm.
Original comment of Harzhauser:
- "The widespread Strombus preoccupatus Finlay 1927 from the Indo-Pacific and western Pacific Miocene is highly reminiscent of Strombus bernielandaui and might represent a closely related species. Strombus preoccupatus has been described by Martin (1881, 1899, 1921) from the Miocene of Java and Borneo, as Strombus spinosus, a name which was already preoccupied as pointed out by Finlay (1927). Strombus bernielandaui differs from Strombus preoccupatus by a shorter and more conical bodywhorl due to the absence of the broad, flat and subparallel area between the shoulder and the first row of spines on the bodywhorl typical of Strombus preoccupatus. Strongly pointed spines on the bodywhorl of Strombus preoccupatus are missing in the new species which develops 2 rows of low and axially elongated knobs. The strongly thickened outer lip is distinctly separated as varix-like swelling from the wing but is developing continuously from the bodywhorl in Strombus preoccupatus. Finally, Strombus preoccupatus develops lirae in the outer lip whilst the Omanian shells display smooth inner lips. Strombus quilonensis Dey 1961 from the (late?) Miocene of Kerala in southern India is also similar but lacks the typical axially elongated knobs on the longer bodywhorl and differs in the strong lirae along the inner lip. It might simply be a less ornamented subspecies of Strombus preoccupatus. This small-sized Strombus is extremely frequent in the Warak Formation at Gebel Madrakah, where it co-occurs with Acropora branches and various cerithiids. In the overlying Ghubbarah Formation it is fully replaced by Strombus gijskronenbergi. Strombus bernielandaui is unknown from the northern coast of the Oligocene Tethys and seems to be a typical southern Tethys species. Its virtually close relation with Strombus preoccupatus suggests that it is a Proto-Indo-Pacific element. The morphologic characters suggest that the Oligocene Strombus bernielandaui might have been the ancestor of the Aquitanian Strombus gijskronenbergi and the Early to Late Miocene Strombus preoccupatus. The latter seems to be followed by Strombus quilonensis. This purely Indo-Pacific lineage might be rooted in the Oligocene to Miocene Strombus radix-bonelli group of the Western Tethys (pers. comm. Gijs Kronenberg). Therefore, this Tethyan strombid-lineage was not only invading the western Atlantic bioprovince, as documented by Lozouet & Maestrati (1986) and Jung & Heintz, 2001, but has also infiltrated the eastern faunas."
- Harzhauser, 2007
- Jung & Heintz, 2001
- Lozouet, P. & Maestrati, P. (1986): Le Strombus granulatus Swainson, 1822 une relique Mesogeenne. Xenophora, Bull. Assoc. Franc. Conchyliol., 31: 11-15.